Darwin accepted blending inheritance, but Fleeming Jenkin calculated that as it mixed traits, natural selection could not accumulate useful traits.

承认混合遗传，但弗莱明•杰肯认为当性状是混合的情况下，自然不能积有益性状。

Haldane applied mathematical analysis to real world examples of natural selection such as the evolution of industrial melanism in peppered moths.

S.霍丹发表了一系列的论，将数学分析应用到真实世界的自然子，如椒粉蛾工业黑变病的进化。