Male-sterility mainly results from the abnormality of mesospore,tapetum tissue and vascular bundle.

中层织、绒毡层织及药隔维管束异常均是雄性败育的因。

Ubisch bodies and lipid bodies are found in the glandular tapetums. In meiosis of microsporocyte, cytokinesisis is a simultaneous type, which produces an isobilateral tetrad.

小孢子母细胞减数分裂胞质分裂为同时型,产生正四面体型四分体。